Punctuation set to "No action (do not replace)" is ignored in favour of separator character

I created a content type called "wiki page", which was configured in Pathauto to follow the following pattern: "wiki/[title-raw]". When I create a wiki page with the title "Test of Wikitools - Pathauto (test page)", my choice of "No action (do not replace)" under punctuation for hyphens, underscores, and both parenthesis is ignored. The alias Pathauto creates is "wiki/Test_of_Wikitools_Pathauto_test_page". As you can deduce, my separator character is the underscore.

I've disabled the wikitools module, since it does hook into the URL callbacks, and I reconfirmed all my settings where in order for Pathauto. My tests returned the same results, with the complete elimination of both the hyphen and extra space, as well as both parenthesis.